The use of colours in this level really make the environment stand out and give it a sense of danger. Having flat coloured textures coupled with the lower poly models makes the scene incredibly bold.
Even in a completely different setting, these chunky models and bold colours keep within a very specific style. Additionally, the character models of both Zack and Wiki have thick black borders around them. While this is present with some of the environment, it lends to make certain parts of the game stand out more.
Thick black borders are also quite a big thing for many cel shaded games such as 'The Legend of Zelda: The Wind Waker', 'Borderlands', and 'Eternal Sonata'. Each of these have incredibly beautiful art styles but don't necessarily fit under the same category as Zack & Wiki. It's evident that it's not just the textures and art style (while this does seem to be a major contributor), that turn a game into one with charm.
More thought, I think, is needed here.
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